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The cellular furin levels were reduced by smoking [331], whilst TMPRSS4 and TMPRSS2 increased and unchanged in bronchial epithelial cells from current smokers in comparison with never smokers [330]

A 90-day nose-only inhalation exposure showed that IQOS had less impact than CS on body weight, hyperplasia and squamous metaplasia in the upper airway, lung inflammation, and overall biological impact (assessed by transcriptomic analysis)